k1 = K1 k2 = K2 k3 = K3 k4 = K4 k5 = Kc k6 = Kd k7 = kd k8 = V2 k9 = V4 k10 = vs k11 = vd k12 = Vm1 k13 = Vm3 k14 = V1 k15 = V3 k16 = compartment