k1 = r_1
k2 = alpha_1
k3 = alpha_2
k4 = k_1
k5 = r_2
k6 = alpha_3
k7 = k_2
k8 = gamma_1
k9 = k_3
k10 = mu_1
k11 = alpha_4
k12 = k_4
k13 = compartment