k1 = alpha k2 = alpha_0 k3 = alpha_1 k4 = delta_2 k5 = mu_1 k6 = mu_10 k7 = mu_11 k8 = mu_2 k9 = mu_20 k10 = beta_1 k11 = beta_2 k12 = mu_3 k13 = mu_30 k14 = gamma_1 k15 = gamma_2 k16 = mu_4 k17 = mu_40 k18 = delta_1 k19 = compartment