k1 = mu_T k2 = r_T k3 = a_1 k4 = b_1 k5 = a_2 k6 = b_2 k7 = mu_G k8 = q_G k9 = p_G k10 = a_G k11 = b_G k12 = mu_I k13 = q_I k14 = p_I k15 = a_I k16 = b_I k17 = c_1 k18 = d_1 k19 = d_2 k20 = c_2 k21 = c_G k22 = d_G k23 = e_G k24 = f_G k25 = L k26 = T_1_e5 k27 = T_2_e5 k28 = compartment